The struggle of God Eaters, like that of the members ofBlood, had ended. The "Devouring Apocalypse", a threat that would engulf the world, changed its form into a structure known as the "Spiral Tree" due to the sacrifice of a God Arcuser.Then, one day something odd in the "Spiral Tree" happens. The members of Blood set out for an investigation together with fellow God Arc user, Livie Collete dispatched by Fenrirheadquarters. While each holds their own feelings in their hearts, they will have to confront with the Spiral Tree once more